Lab Publications


Bibi F, Pante M, Souron A, Stewart K, Varela S, Werdelin L, Boisserie J-R, Fortelius M, Hlusko L, Njau J, de la Torre I. In press. Paleoecology of the Serengeti during the Oldowan-Acheulean transition at Olduvai Gorge, Tanzania: The mammal and fish evidence. Journal of Human Evolution

Reiner WB, Masao F, Sholts S, Songita A, Stanistreet I, Stollhofen H, Taylor RE, Hlusko LJ. In press. OH 83: A new early modern human fossil cranium from the Ndutu Beds of Olduvai Gorge, Tanzania. American Journal of Physical Anthropology available in early view DOI 10.1002/ajpa.23292

Monson TA, Brasil MF, Stratford DJ, Hlusko LJ. 2017. Patterns of craniofacial variation and taxonomic diversity in the South African Cercopithecidae fossil record. Palaeontologia Electronica 20.1.7A: 1-20


Monson TA, Hlusko LJ. 2016. The evolution of dental eruption sequence in artiodactyls. Journal of Mammalian Evolution DOI 10.1007/s10914-016-9362-9 published online 28 Nov 2016

Hlusko LJ, Schmitt CA, Monson TA, Brasil MF, Mahaney MC. 2016. The integration of quantitative genetics, paleontology, and neontology reveals genetic underpinnings of primate dental evolution. Proceedings of the National Academy of Sciences USA 113 (33) 92629267 (open access)

Commentary: P. David Polly. Quantitative genetics provides predictive power for paleontological studies of morphological evolution. Proceedings of the National Academcy of Sciences USA 113(33): 9142-9144. 

Ungar PS, Hlusko LJ. 2016. The evolutionary path of least resistance. Science 353(6294):29-30.  (2016_Science)

Grieco TM, Hlusko LJ. 2016. Insight from frogs: Sonic hedgehog gene expression and a re-evaluation of the vertebrate odontogenic band. The Anatomical Record 299(8):1099-109.  (2016_anatomical_record)

Hlusko LJ. 2016. Elucidating the evolution of the hominid dentition in the age of phenomics, modularity, and quantitative genetics. Annals of Anatomy 203:3-11.  (2016_Annals_Anatomy)


Lainoff AJ, Mousakas JE, Kallonen A, Hämälainen K, Smith KK, Hu D, Marcucio R, Hlusko LJ. 2015. A comparative examination of odontogenic markers in both toothed and toothless amniotes. Molecular and Developmental Evolution (J. Exp. Zoolog.) 324(3):255-69. (2015_J_ExpZool_Mol_Dev_Evol)

Monson TA, Brasil MF, Hlusko LJ. 2015. Materials collected by the southern branch of the UC Africa Expedition with a report on new Plio-Pleistocene fossil localities. PaleoBios 32. ucmp_paleobios_29407. (2015_PaleoBios)

Hlusko LJ, Reiner WR, Njau JK. 2015. A one-million-year-old hominid distal ulna from Olduvai Gorge, Tanzania. American Journal of Physical Anthropology 158(1):36-42.  (2015_Am_J_Phys_Anth)


Reiner WB, Petzinger C, Power ML, Hyeroba D and JM Rothman. 2014. Fatty acids in mountain gorilla diets: implications for primate nutrition and health. American Journal of Primatology 76(3):281-8.

Monson T, Hlusko LJ. 2014. Identification of a derived dental trait in the Papionini relative to other Old World Monkeys.American Journal of Physical Anthropology 155:422-429.  (2014_Am_J_Phys_Anth)

Amugongo SK, Hlusko LJ. 2014.Impact of maternal prenatal stress on growth of the offspring. Aging and Disease 5(1):1-16.  (2014_Aging_and_Disease)

Morrish KR, Hlusko LJ. 2014. Modularity and sexual dimorphism in human metacarpals. Paleobios 31(2):1-20.  (2014_Paleobios)


evo_67_1_coverGrieco TM, OT Rizk, LJ Hlusko. 2013. A modular framework characterizes micro- and macroevolution of Old World monkey dentitions. Evolution 67(1):241-59. (2013_Evolution)

Hlusko LJ, Carlson J, Guatelli-Steinberg D, Krueger KL, Mersey B, Ungar PS, Defleur A. 2013. Neanderthal teeth from Moula-Guercy, Ardeche, France. American Journal of Physical Anthropology 151:477-491.  (2013_Am_J_Phys_Anth)

Rizk OT, Grieco TM, Holmes MW, LJ Hlusko. 2013. Using geometric morphometrics to study the mechanisms that pattern primate dental variation. Anthropological Perspectives on Tooth Morphology: Genetics, Evolution, Variation, edited by Dr. G. Richard Scott and Dr. Joel Irish  (2013_GM_chapt_proofs)


Renne PR., Mulcahy SR, Morgan LE, Cassata WS, Kelley SP, Hlusko LJ, Njau JK. 2012. Retention of inherited Ar by alkali feldspar xenocrysts in a magma: Kinetic constraints from Ba zoning profiles. Geochimica et Cosmochimica Acta 93:129-142.  (2012_Geo_Cosmo_Acta)


Moustakas JE, KK Smith, LJ Hlusko. 2011. The evolution and development of the mammalian dentition: insights from the marsupial Monodelphis domestica. Developmental Dynamics 240(1):232-9.  (2011_Dev_Dynamics)

Hlusko LJ, RD Sage, MC Mahaney. 2011. Evolution of modularity in the mammalian dentition: mice and monkeys share a common dental genetic architecture. Molecular and Developmental Evolution (J. Exp. Zoolog.) 316B:21-49.  (2011_J_Exp_Zool_Mol_Dev_Evol)


Grieco TM, OT Rizk. 2010. Cranial shape varies along an elevation gradient in Gambel’s White-Footed mouse (Peromyscus maniculatus gambelii) in the Grinnell Resurvey Yosemite Transect. Journal of Morphology 271:897-909.

Njau JK and Hlusko LJ. 2010. Fine-tuning paleoanthropological reconnaissance using high-resolution satellite imagery: the discovery of 28 new sites in Tanzania. Journal of Human Evolution 59:680-684.  (2010_J_Hum_Evol)

Koh C, Bates E, Broughton E, Do NT, Fletcher Z, Mahaney MC, and Hlusko LJ. 2010. Genetic integration of molar cusp size variation in baboons. American Journal of Physical Anthropology 142:246-260.  (2010_Am_J_Phys_Anth)


Louchart A, Wesselman H, Blumenschine RJ, Hlusko LJ, Njau JK, Black MT, Asnake M, White TD. 2009. Taphonomic, avian, and small vertebrate indicators of Ardipithecus ramidus habitat. Science 326:66 (66e1-66e4)  (2009_Science)

Hlusko LJ and MC Mahaney. 2009. Quantitative genetics, pleiotropy, and morphological integration in the dentition of Papio hamadryas. Evolutionary Biology 36(1):5 (DOI 10.1007/s11692-008-9048-1)  (2009_Evol_Biol)

Hlusko LJ and Mahaney MC. 2009. Baboon model for dental development. In: The Baboon in Biomedical Research, Southwest Foundation for Biomedical Research Press, San Antonio, TX. Springer, New York. Pages 207-223.  (2009_Bab_Biomed_Res)

Frost SR, Haile-Selassie Y, and Hlusko LJ. 2009. Chapter 6 Cercopithecidae. In: Ardipithecus kadabba: Late Miocene Evidence from Middle Awash, Ethiopia (Y. Haile-Selassie, G WoldeGabriel eds). University of California Press, Berkeley.


Boisserie, J-R, Guy F, Delagnes A, Hlusko LJ, Bibi F, Beyene Y, and Guillemot C. 2008. New paleoanthropological research in the Plio-Pleistocene Omo Group, Lower Omo Valley, SNNPR (Southern Nations, Nationalities and People Regions), Ethiopia. C. R. Palevol 7:429-439.  (2008_Palevol)

Rizk OT, Amugongo S, Mahaney MC, and Hlusko LJ. 2008. The quantitative genetic analysis of primate dental variation: History of the approach and prospects for the future. In: Technique and Application in Dental Anthropology (J.D. Irish and G.C. Nelson, eds). Cambridge University Press. (ISBN-13:9780521870610).  (2008_rizk_etal_chapt)

Hlusko LJ. 2008. Book Review. The Ape in the Tree: An Intellectual and Natural History of Proconsul, by A. Walker and P. Shipman. Current Anthropology 49(2):337-338.  (2008_Curr_Anth_book_review)


For the complete Lemudon’go monograph, please go to this link.

Hlusko LJ and Mahaney MC. 2007. Of mice and monkeys: Quantitative genetic analyses of size variation along the dental arcade. In: Dental Perspectives on Human Evolution: State of the Art Research in Dental Paleoanthropology (S. Bailey and J-J. Hublin, eds.). Springer, Dordrecht, pp. 237-245.  (2007_Dental_Perspectives)

Hlusko LJ, Do N, Mahaney MC. 2007. Genetic correlations between mandibular molar cusp areas in baboons. American Journal of Physical Anthropology 132:445-454.  (2007_Am_J_Phys_Anth)

Hlusko LJ and Mahaney MC. 2007. A multivariate comparison of dental variation in wild and captive populations of baboons (Papio hamadryas). Archives of Oral Biology 52:195-200.  (2007_Arch_Oral_Biol)

Hlusko LJ. 2007. A new late Miocene species of Paracolobus and other Cercopithecoidea (Mammalia: Primates) fossils from Lemudong’o, Kenya. Kirtlandia 56: 72-85.  (2007_Kirtlandia_paracolobus)

Hlusko LJ. 2007. Earliest evidence for Atherurus and Xenohystrix (Hystricidae, Rodentia) in Africa, from the late Miocene site of Lemudong’o, Kenya. Kirtlandia 56: 86-91.

Hlusko LJ and Y Haile-Selassie. 2007. Nyanzachoerus syrticus (Artiodactyla, Suidae) from the late Miocene of Lemudong’o, Kenya. Kirtlandia 56: 152-157.

Hlusko LJ, Y Haile-Selassie, D Degusta. 2007. Late Miocene Bovidae (Mammalia: Artiodactyla) from Lemudong’o, Narok District, Kenya. Kirtlandia 56: 163-172.

Ambrose SH, Bell CJ, Bernor RL, Boisserie JR, Darwent CM, DeGusta D, Deino A, Garcia N, Haile-Selassie Y, Head JJ, Howell FC, Kyule MD, Manthi FK, Mathu EM, Nyamai CM, Pickford M, Saegusa H, Stidham TA, Williams MAJ, and Hlusko LJ. 2007. The paleoecology and paleogeographic context of Lemudong’o Locality 1, a Late Miocene terrestrial fossil site in Southern Kenya. Kirtlandia 56:38-52.  (2007_Kirtlandia_paleoecology)

Ambrose SH, MD Kyule, LJ Hlusko. 2007. History of paleontological research in the Narok District of Kenya. Kirtlandia 56: 1-37.  (2007_Kirtlandia_history)

Saegusa H and LJ Hlusko. 2007. New late Miocene elephantoid (Mammalia: Proboscidea) fossils from Lemudong’o, Kenya. Kirtlandia 56: 140-147.

Pickford M and LJ Hlusko. 2007. Late Miocene Procaviid hyracoids (Hyracoidea: Dendrohyrax) from Lemudong’o, Kenya. Kirtlandia 56: 106-111.


Hlusko LJ, Lease LR, Mahaney MC. 2006. The evolution of genetically correlated traits: tooth size and body size in baboons.American Journal of Physical Anthropology 131:420-427.  (2006_Am_J_Phys_Anth)

White TD, WoldeGabriel G, Asfaw B, Ambrose S, Beyene Y, Bernor RL, Boisserie JR, Currie B, Gilbert H, Haile-Selassie Y, Hart WK, Hlusko LJ, Howell FC, Kono RT, Louchart A, Lovejoy CO, Renne PR, Saegusa H, Vrba E, Wesselman H, and Suwa G. 2006. Asa Issie, Aramis, and the origin of Australopithecus. Nature 440:883-889.

Hlusko LJ. 2006. A New Large Pliocene Colobine Species (Mammalia: Primates) from Asa Issie, Ethiopia. Geobios 39:57-69.  (2006_Geobios)

Hlusko LJ. 2006. Book Review: The Human Fossil Record, Volume Four: Craniodental Morphology of Early Hominids (General Australopithecus, Paranthropus, Orrorin), and Overview by J.H. Schwartz and I. Tattersall. American Journal of Physical Anthropology 131:148-149.  (2006_Am_J_Phys_Anth_book_review)


Sherwood RJ, Hlusko LJ, Duren DL, Ench VC, and Walker A. 2005. The mandibular symphysis of large-bodied hominoids. Human Biology 77(6):735-759. (2005_Hum_Bio)

Haile-Selassie Y, Hlusko LJ, and Howell FC. 2004. A new species of Plesiogulo (Mustelidae, Carnivora) from the Late Miocene of Africa. Palaeontologia Africana 40:85-88.  (2004_Pal_afr)


Hlusko LJ, Maas ML, Mahaney MC. 2004. Statistical genetics of molar cusp patterning in pedigreed baboons: Implications for primate dental development and evolution. Molecular and Developmental Evolution (J. Exp. Zoolog.) 302B:268-283. (2004_Mol_Dev_Evol)

Hlusko LJ. 2004. Protostylid variation in Australopithecus. Journal of Human Evolution 46: 579-594.  (2004_J_Hum_Evol)

Hlusko LJ. 2004. Integrating the genotype and phenotype in hominid paleontology. Proceedings of the National Academy of Sciences USA 101(9): 2653-2657.  (2004_Proc_Natl_Acad_Sci)

Hlusko LJ, Suwa G, Kono R, and Mahaney MC. 2004. Genetics and the evolution of primate enamel thickness: A baboon model.American Journal of Physical Anthropology 124:223-233.   (2004_Am_J_Phys_Anth)


Hlusko LJ. 2003. The oldest hominid habit? Experimental evidence for toothpicking with grass stalks. Current Anthropology44(5): 738-741.  (2003_Curr_Anth)

Hlusko LJ and Mahaney MC. 2003. Genetic contributions to expression of the baboon cingular remnant. Archives of Oral Biology 48: 663-672.  (2003_Arch_Oral_Biol)

Ambrose SH, Hlusko LJ, Kyule D, Deino A, and Williams M. 2003. Lemudong’o: A new 6ma paleontological site near Narok, Kenya Rift Valley. Journal of Human Evolution 44:737-742.  (2003_J_Hum_Evol)

Hlusko LJ. 2003. Book Review: Colbert’s Evolution of the Vertebrates by E.H. Colbert, M. Morales, and E.C. Minkoff. American Journal of Physical Anthropology 123:97.  (2003_Am_J_Phys_Anth_book_review)


Hlusko LJ. 2002. Identifying metameric variation in extant hominoid and fossil hominid molars. American Journal of Physical Anthropology 118(1): 86-97.  (2002a_Am_J_Phys_Anth)

Hlusko LJ. 2002. Expression types for two cercopithecoid dental traits (Interconulus and Interconulid) and their variation in a modern baboon population. International Journal of Primatology 23(6): 1309-1318.  (2002_Intl_J_Primatol)

Hlusko LJ, Mahaney MC, and Weiss KM. 2002. A statistical genetic comparison of two techniques for assessing molar crown size in pedigreed baboons. American Journal of Physical Anthropology 117(2): 182-189.  (2002b_Am_J_Phys_Anth)

Hlusko LJ. 2002. Book Review: Extinct Humans, by I. Tattersall and J.H. Schwartz. Quarterly Review of Biology 77(4): 446-447.  (2002_QRB_book_review.web)


Liu W, LJ Hlusko, and Zheng. 2001. Morphometric analysis of hominoid lower molars from Yuanmou of Yunnan Province, China.Primates 42(2): 123-134.  (2001_Primates)


Hlusko LJ. 2000. Identifying the genetic mechanisms of dental variation in cercopithecoid primates. PhD dissertation, The Pennsylvania State University, UMI 9982340.